Sorry I wasn't more clear. Did they provide the causes for the decrease in nutrition in the forage? Change to a lower quality species? Same species, but they are less nutritious than in the past?
Main causes are warmer temps and drier conditions leading to less nutritional value.
They do talk about increased predation on the migratory elk as well, but when you don't get young cows reproducing, nothing matters in terms of predation because you never replenish the herd.
Here's the link to the web page: http://www.wyocoopunit.org/index.php/kauffman-group/search/absaroka-elk-ecology-project/
Nectar- as it relates to this study, it's due to climate change and drought primarily.